A. Gimpel et al.
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Science of the Total Environment 878 (2023) 162902
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Fig. 2. Length frequencies of cod individuals sampled inside the Offshore Wind Farm during summer (OWF summer, yellow) (A); inside the Offshore Wind Farm during
winter (OWF winter, blue) (B); at stations outside the OWF in the German Bight during summer (GB summer, red) (C); at stations outside the OWF in the German Bight
during winter (GB winter, grey) (D). Further given are the lengths at age of cod individuals sampled inside the Offshore Wind Farm during summer (yellow), during
winter (blue) and at stations outside the OWF in the German Bight during summer (red) and winter (grey) (E). Moreover, the relative proportion of individuals from
different Surveys per age group is shown (OWF summer = yellow; OWF winter = blue; GB summer = red and GB winter = grey). Please note: On-site samples
collected by angling were combined with data from outside the OWF collected with bottom trawls. (For interpretation of the references to color in this figure legend, the
reader is referred to the web version of this article.)
ındividuals in the OWF in summer. Nevertheless, the majority of individ-
Jals +3 years were caught in the German Bight (Fig. 2E-F).
We observed spatial differences in stomach contents, with cod caught in
+ he OWF having a more diverse prey composition compared to samples from
he GB (Fig. 3). A CCA was used to investigate the relationship between body
length, study area and diet composition (Fig. 4). Both predictors and their in-
:eraction were highly significant (Table 2), indicating that diet composition
changes ontogenetically and differs between study areas. Moreover, the im-
pact of the study area on diet varies with size class, as indicated by the signif-
icant interactive term (Fig. 3). The diet of cod caught in the GB consisted of
flatfish and brown shrimp (Crangon sp.), which are typically associated to
sandy bottoms (Fig. 4, Appendix A, Table A.1). In contrast, cod in the
JWF fed mainly on species associated to hard substrates, such as porcelain
crab or butterfish (Fig. 4). Fish above 45 cm were associated with a diet
zonsisting of flatfish and swimming crabs, while the diet of smaller fish
was associated with consumption of shrimps, for instance (Fig. 4).
individuals selected for stable isotope analyses confirmed the observed
variation of stomach contents between the OWF and the GB samples. The
1igh 8*°N results in the OWF in summer suggest that cod fed higher in
:he food web than either in GB summer or in OWF winter (Fig. 5,
Table 1). Mean 8'°N differed significantly between cod from stations sam-
led in the GB with 15.37 %o (+ 1.49) and cod sampled in the OWF during